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MammalBase - Accepted taxon: Naemorhedus goral

Accepted taxon: Naemorhedus goral


Accepted taxon: Naemorhedus goral
Taxon: Naemorhedus goral
Rank: Species
Higher Classification: ARTIODACTYLA | Bovidae | Caprinae | Naemorhedus
Reference: Wilson, Don E., and DeeAnn M. Reeder, eds. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), JHU Press, 2005.

Fig.1 - Ternary plot of the proximate analysis for the Naemorhedus goral diet.
CF (Crude Fibre) 27, ASH (Ash contents) 9, NFE (Nitrogen-Free-Extract) 50, CP (Crude Protein) 11, EE (Ether Extract) 3. See details for the calculations in Lintulaakso, K., Tatti, N. and Žliobaitė, I., 2022. Quantifying mammalian diets. Mammalian Biology, pp.1-15. [DOI]


Table 1. List of Diet Items for Naemorhedus goral.
TSN Complete name Part Ratio a. Reference(s)
Poaceae STEM 45.00 Grzimek,B…
Magnoliopsida SHOOT 31.67 Grzimek,B…
Magnoliopsida STEM 10.00 Smith,A. …
Magnoliopsida LEAF 8.33 Grzimek,B…
Magnoliopsida FRUIT 5.00 Smith,A. …

a. Standardized values. See details for the calculations in Lintulaakso, K., Tatti, N. and Žliobaitė, I., 2022. Quantifying mammalian diets. Mammalian Biology, pp.1-15. [DOI]


Table 2. List of Standard Traits for Naemorhedus goral.
Trait Selected Value All Value(s) Reference(s)
Dietary category: Eisenberg 1981 Herbivore-Mixed feeder Herbivore-Browser, Herbivore-Grazer, Herbivore-Mixed feeder Abraham, J.O., Upham, N.S., Damian-Serrano, A. and Jesmer, B.R., 2022. Evolutionary causes and consequences of ungulate migration. Nature Ecology & Evolution, 6(7), pp.998-1006., I. J. Gordon, A. W. Illius, 1988, 'Incisor Arcade Structure and Diet Selection in Ruminants', Functional Ecology, vol. 2, no. 1, p. 15, Loison,A., 1999, What factors shape sexual size dimorphism in ungulates?, Evolutionary Ecology Research, M. Mendoza, P. Palmqvist, 2008, 'Hypsodonty in ungulates: an adaptation for grass consumption or for foraging in open habitat?', Journal of Zoology, vol. 274, no. 2, pp. 134-142, Marcus Clauss, Matthias Lechner-Doll, Juergen Streich, 2002, 'Faecal particle size distribution in captive wild ruminants: an approach to the browser/grazer dichotomy from the other end', Oecologia, vol. 131, no. 3, pp. 343-349, Quibod, M., Gélin, U., van Langevelde, F., Tomlinson, K. 2023. Diet-specific responses of skull traits to aridity gradients in bovids and cervids. Zoological Journal of the Linnean Society. None(None), pp.None. Available at: 10.1093/zoolinnean/zlad068.
Dietary category: Miljutin 2009 Herbivore Herbivore Abraham, J.O., Upham, N.S., Damian-Serrano, A. and Jesmer, B.R., 2022. Evolutionary causes and consequences of ungulate migration. Nature Ecology & Evolution, 6(7), pp.998-1006., I. J. Gordon, A. W. Illius, 1988, 'Incisor Arcade Structure and Diet Selection in Ruminants', Functional Ecology, vol. 2, no. 1, p. 15, Loison,A., 1999, What factors shape sexual size dimorphism in ungulates?, Evolutionary Ecology Research, M. Mendoza, P. Palmqvist, 2008, 'Hypsodonty in ungulates: an adaptation for grass consumption or for foraging in open habitat?', Journal of Zoology, vol. 274, no. 2, pp. 134-142, Marcus Clauss, Matthias Lechner-Doll, Juergen Streich, 2002, 'Faecal particle size distribution in captive wild ruminants: an approach to the browser/grazer dichotomy from the other end', Oecologia, vol. 131, no. 3, pp. 343-349, Quibod, M., Gélin, U., van Langevelde, F., Tomlinson, K. 2023. Diet-specific responses of skull traits to aridity gradients in bovids and cervids. Zoological Journal of the Linnean Society. None(None), pp.None. Available at: 10.1093/zoolinnean/zlad068.

Table 3. List of Standard Measurements for Naemorhedus goral.
Measurement Location Unit n Total n Unknown n Female n Male Min Mean Max SD Reference(s)
Body Mass (BM) nan g - - - - - 29,360 - - Wimberly,…
Body Mass (BM) nan g - - - - 22,000 - 42,000 - Freudenth…

Table 4. List of other Measurements b. for Naemorhedus goral.
Measurement Location Unit n Total n Unknown n Female n Male Min Mean Max SD Reference(s)
Length of the Masseteric Fossa cm 6 6 - - - 11 - - Quibod, M…
Muzzle Width cm 6 6 - - - 2 - - Quibod, M…
Total Length of the Skull cm 6 6 - - - 26 - - Quibod, M…
A1-MLL (maximum lateral length of the astragalus) nan mm 6 6 - - 28 31 34 2 Wimberly,…
A2-MML (maximum medial length of the astragalus) nan mm 6 6 - - 27 29 32 2 Wimberly,…
A3-MIN (minimal length of the astragalus) nan mm 6 6 - - 23 25 28 2 Wimberly,…
A4-WAT (width at tarsus of the astragalus) nan mm 6 6 - - 17 20 22 2 Wimberly,…
A5-WAF (width at trochlea of the astragalus) nan mm 6 6 - - 15 18 19 2 Wimberly,…
C1-UL (greatest length of the calcaneus) nan mm 6 6 - - 59 63 69 5 Wimberly,…
C2-TL (length of body of the calcaneus) nan mm 6 6 - - 39 42 44 2 Wimberly,…
C3-MBH (midbody height of the calcaneus) nan mm 6 6 - - 14 16 17 1 Wimberly,…
C4-MBTV (midbody transverse width of the calcaneus) nan mm 6 6 - - 7 8 9 1 Wimberly,…
C5-FFL (fibular facet length of the calcaneus) nan mm 6 6 - - 12 13 14 1 Wimberly,…
C6-CNFL (cubonavicular facet length of the calcaneus) nan mm 6 6 - - 14 17 18 2 Wimberly,…
C7-HSF (height of sustentacular facet of the calcaneus) nan mm 6 6 - - 13 14 15 1 Wimberly,…
C8-HST (height of sustentaculum tali of the calcaneus) nan mm 6 6 - - 19 20 22 2 Wimberly,…
F1-UL (full length of the femur) nan mm 6 6 - - 183 193 213 11 Wimberly,…
F2-FL (functional length of the femur) nan mm 6 6 - - 178 190 207 11 Wimberly,…
F3-MCF (midshaft circumference of the femur) nan mm 6 6 - - 48 54 61 4 Wimberly,…
F4-PW (proximal width of the femur) nan mm 6 6 - - 37 43 46 3 Wimberly,…
F5-MLHW (mediolateral femoral head of the femur) nan mm 6 6 - - 21 24 26 2 Wimberly,…
F6-APHW (anteroposterior femoral head of the femur) nan mm 6 6 - - 18 21 22 2 Wimberly,…
F7-DW (distal width of the femur) nan mm 6 6 - - 35 39 44 3 Wimberly,…
F8-MPL (medial condyle and patellar lip of the femur) nan mm 6 6 - - 40 44 47 3 Wimberly,…
F9-LPL (lateral condyle and patellar lip of the femur) nan mm 6 6 - - 39 41 43 2 Wimberly,…
H1-UL (full length of the humerus) nan mm 6 6 - - 176 184 199 10 Wimberly,…
H2-FL (functional length of the humerus) nan mm 6 6 - - 160 168 183 10 Wimberly,…
H3-MCF (midshaft circumference of the humerus) nan mm 6 6 - - 52 57 66 6 Wimberly,…
H4-MTH (largest medial height of trochlea of the humerus) nan mm 6 6 - - 18 19 20 1 Wimberly,…
H5-MidTH (smallest height of trochlea of the humerus) nan mm 6 6 - - 14 15 16 1 Wimberly,…
H6-LTH (largest lateral height of trochlea of the humerus) nan mm 6 6 - - 16 17 18 1 Wimberly,…
H7-PW (proximal width of the humerus) nan mm 6 6 - - 38 40 43 2 Wimberly,…
H8-MLH (mediolateral humeral head width of the humerus) nan mm 6 6 - - 30 31 33 1 Wimberly,…
H9-DW (distal width of the humerus) nan mm 6 6 - - 31 33 37 3 Wimberly,…
MC1-UL (length of the metacarpus) nan mm 6 6 - - 111 121 127 7 Wimberly,…
MC2-MAP (anteroposterior midshaft of the metacarpus) nan mm 6 6 - - 12 12 13 0 Wimberly,…
MC3-MML (mediolateral midshaft of the metacarpus) nan mm 6 6 - - 17 18 19 1 Wimberly,…
MC4-DW (distal width of the metacarpus) nan mm 6 6 - - 27 29 30 1 Wimberly,…
MC5-PW (proximal width of the metacarpus) nan mm 6 6 - - 26 27 28 1 Wimberly,…
MT1-UL (length of the metatarsus) nan mm 6 6 - - 118 126 132 6 Wimberly,…
MT2-MAP (anteroposterior midshaft of the metatarsus) nan mm 6 6 - - 12 13 14 1 Wimberly,…
MT3-MML (mediolateral midshaft of the metatarsus) nan mm 6 6 - - 12 14 15 1 Wimberly,…
MT4-DW (distal width of the metatarsus) nan mm 6 6 - - 24 26 27 1 Wimberly,…
MT5-PW (proximal width of the metatarsus) nan mm 6 6 - - 21 23 24 2 Wimberly,…
R1-UL (length of the radius) nan mm 6 6 - - 157 162 174 7 Wimberly,…
R2-PW (full proximal width of the radius) nan mm 6 6 - - 28 30 33 2 Wimberly,…
R3-NPW (width of humeral articular surface of the radius) nan mm 6 6 - - 28 30 32 2 Wimberly,…
R4-DW (distal width of the radius) nan mm 6 6 - - 27 30 33 3 Wimberly,…
S1-L (full length of the scapula) nan mm 6 6 - - 135 140 145 5 Wimberly,…
S2-GW (greatest cranio-caudal length of the scapula) nan mm 6 6 - - 75 85 90 8 Wimberly,…
S3-SW (shortest length of scapular neck of the scapula) nan mm 6 6 - - 17 18 19 1 Wimberly,…
S4-GH (anteroposterior glenoid fossa of the scapula) nan mm 6 6 - - 22 24 25 1 Wimberly,…
S5-GW (mediolateral glenoid fossa of the scapula) nan mm 6 6 - - 20 22 23 1 Wimberly,…
T1-L (full length of the tibia) nan mm 6 6 - - 192 222 238 16 Wimberly,…
T2-MCF (midshaft circumference of the tibia) nan mm 6 6 - - 45 50 56 4 Wimberly,…
T3-PW (proximal width of the tibia) nan mm 6 6 - - 37 41 46 4 Wimberly,…
T4-DW (distal width of the tibia) nan mm 6 6 - - 25 28 31 2 Wimberly,…
U1-UL (length of the ulna) nan mm 6 6 - - 201 209 226 10 Wimberly,…
U2-OPL (olecranon process length of the ulna) nan mm 6 6 - - 39 42 46 3 Wimberly,…
average ungulate dietary grass fraction (proportional) nan unitless - - - - - 1 - - Abraham, …
femur diameter nan mm - - - - - 17 - - Nelson, A…
humerus diameter nan mm - - - - - 17 - - Nelson, A…
neck length nan mm - - - - - 213 - - Nelson, A…

b. other Measurements. Source Measurements which are not listed on or mapped with the Standard Traits..


Table 5. List of Data Sources for Naemorhedus goral.
Taxon Reference Match Status
Nemorhoedes goral Gordon,IJ, 1988, Incisor arcade structure and diet selection in ruminants, Functional Ecology 150: Genus synonym, same species epithet Verified - Accepted
Nemorhaedus goral Loison,A., 1999, What factors shape sexual size dimorphism in ungulates?, Evolutionary Ecology Research 51: Name variant or misspelling of the taxon Verified - Accepted
Nemorhaedus goral Clauss,M., 2002, Faecal particle size distribution in captive wild ruminants: an approach to the browser/grazer dichotomy from the other end, Oecologia 51: Name variant or misspelling of the taxon Verified - Accepted
Nemorhaedus goral Mendoza,M., 2008, Hypsodonty in ungulates: an adaptation for grass consumption or for foraging in open habitat?, Journal of zoology 51: Name variant or misspelling of the taxon Verified - Accepted
Naemorhedus goral Grzimek,B., 2003, Grzimek's Animal Life Encyclopedia: Mammals I-V, 1: Exact Taxon match Verified - Accepted
Naemorhedus goral Smith,A. T., 2010, A guide to the mammals of China, 1: Exact Taxon match Verified - Accepted
Naemorhedus goral Quibod, M.N.R.M., Gélin, U., van Langevelde, F. and Tomlinson, K.W., 2023. Diet-specific responses of skull traits to aridity gradients in bovids and cervids. Zoological Journal of the Linnean Society, p.zlad068. 1: Exact Taxon match Verified - Accepted
Naemorhedus goral Wimberly, A.N., 2023. Predicting body mass in Ruminantia using postcranial measurements. Journal of Morphology, 284(10), p.e21636. 1: Exact Taxon match Verified - Accepted
Naemorhedus goral Abraham, J.O., Upham, N.S., Damian-Serrano, A. and Jesmer, B.R., 2022. Evolutionary causes and consequences of ungulate migration. Nature Ecology & Evolution, 6(7), pp.998-1006. 1: Exact Taxon match Verified - Accepted
Naemorhedus goral Lundgren, E.J., Schowanek, S.D., Rowan, J., Middleton, O., Pedersen, R.Ø., Wallach, A.D., Ramp, D., Davis, M., Sandom, C.J. and Svenning, J.C., 2021. Functional traits of the world’s late Quaternary large-bodied avian and mammalian herbivores. Scientific data, 8(1), p.17. 1: Exact Taxon match Verified - Accepted
Naemorhedus goral Nelson, A., Engelman, R.K. and Croft, D.A., 2023. How to weigh a fossil mammal? South American notoungulates as a case study for estimating body mass in extinct clades. Journal of Mammalian Evolution, 30(3), pp.773-809. 1: Exact Taxon match Verified - Accepted
Naemorhedus goral Freudenthal, M. and Martín-Suárez, E., 2013. Estimating body mass of fossil rodents. Scripta Geologica, 145, pp.1-513. 1: Exact Taxon match Verified - Accepted
Naemorhedus goral Rowan, J., Beaudrot, L., Franklin, J., Reed, K.E., Smail, I.E., Zamora, A. and Kamilar, J.M., 2020. Geographically divergent evolutionary and ecological legacies shape mammal biodiversity in the global tropics and subtropics. Proceedings of the National Academy of Sciences, 117(3), pp.1559-1565. 1: Exact Taxon match Verified - Accepted

Citation for this page: [The MammalBase community 2026. / CC BY 4.0. http://doi.org/10.5281/zenodo.7462864 Naemorhedus goral , Accessed 1st May 2026 at https://mammalbase.net/me/16089]