MammalBase - Accepted taxon: Capricornis sumatraensis

Accepted taxon: Capricornis sumatraensis
Taxon:	Capricornis sumatraensis
Rank:	Species
Higher Classification:	ARTIODACTYLA \| Bovidae \| Caprinae \| Capricornis
Reference:	Wilson, Don E., and DeeAnn M. Reeder, eds. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), JHU Press, 2005.

Fig.1 - Ternary plot of the proximate analysis for the Capricornis sumatraensis diet.
CF (Crude Fibre) 28, ASH (Ash contents) 9, NFE (Nitrogen-Free-Extract) 49, CP (Crude Protein) 11, EE (Ether Extract) 3. See details for the calculations in Lintulaakso, K., Tatti, N. and Žliobaitė, I., 2022. Quantifying mammalian diets. Mammalian Biology, pp.1-15. [DOI]

Table 1. List of Diet Items for Capricornis sumatraensis.
TSN Complete name	Part	Ratio ^a.	Reference(s)
Gramineae	STEM	29.64	Grzimek,B…
Magnoliopsida	SHOOT	19.06	Grzimek,B…
Magnoliopsida	LEAF	15.71	Grzimek,B…
Magnoliopsida	STEM	12.24	Quyoom, I…
Quercus	LEAF	7.14	Quyoom, I…
Arundinaria	STEM	4.59	Quyoom, I…
Gaultheria	LEAF	3.57	Quyoom, I…
Urtica	SHOOT	3.06	Quyoom, I…
Strobilanthes	SHOOT	2.30	Quyoom, I…
Coniferophyta	LEAF	1.90	Quyoom, I…
Rhododendron	LEAF	0.77	Quyoom, I…

^a. Standardized values. See details for the calculations in Lintulaakso, K., Tatti, N. and Žliobaitė, I., 2022. Quantifying mammalian diets. Mammalian Biology, pp.1-15. [DOI]

Table 2. List of Standard Traits for Capricornis sumatraensis.
Trait	Selected Value	All Value(s)	Reference(s)
Dietary category: Eisenberg 1981	Herbivore-Browser	Herbivore-Browser, Herbivore-Mixed feeder	– Abraham, J.O., Upham, N.S., Damian-Serrano, A. and Jesmer, B.R., 2022. Evolutionary causes and consequences of ungulate migration. Nature Ecology & Evolution, 6(7), pp.998-1006., Loison,A., 1999, What factors shape sexual size dimorphism in ungulates?, Evolutionary Ecology Research, M. Mendoza, P. Palmqvist, 2008, 'Hypsodonty in ungulates: an adaptation for grass consumption or for foraging in open habitat?', Journal of Zoology, vol. 274, no. 2, pp. 134-142, Quibod, M., Gélin, U., van Langevelde, F., Tomlinson, K. 2023. Diet-specific responses of skull traits to aridity gradients in bovids and cervids. Zoological Journal of the Linnean Society. None(None), pp.None. Available at: 10.1093/zoolinnean/zlad068.
Dietary category: Miljutin 2009	Herbivore	Herbivore	– Abraham, J.O., Upham, N.S., Damian-Serrano, A. and Jesmer, B.R., 2022. Evolutionary causes and consequences of ungulate migration. Nature Ecology & Evolution, 6(7), pp.998-1006., Loison,A., 1999, What factors shape sexual size dimorphism in ungulates?, Evolutionary Ecology Research, M. Mendoza, P. Palmqvist, 2008, 'Hypsodonty in ungulates: an adaptation for grass consumption or for foraging in open habitat?', Journal of Zoology, vol. 274, no. 2, pp. 134-142, Quibod, M., Gélin, U., van Langevelde, F., Tomlinson, K. 2023. Diet-specific responses of skull traits to aridity gradients in bovids and cervids. Zoological Journal of the Linnean Society. None(None), pp.None. Available at: 10.1093/zoolinnean/zlad068.

Table 3. List of Standard Measurements for Capricornis sumatraensis.
Measurement	Location	Unit	n Total	n Unknown	n Female	n Male	Min	Mean	Max	SD	Reference(s)
Body Mass (BM)	nan	g	-	-	-	-	75,000	-	140,000	-	Freudenth…
Body Mass (BM)	nan	g	-	-	-	-	-	98,650	-	-	Wimberly,…

Table 4. List of other Measurements ^b. for Capricornis sumatraensis.
Measurement	Location	Unit	n Total	n Unknown	n Female	n Male	Min	Mean	Max	SD	Reference(s)
Length of the Masseteric Fossa		cm	9	9	-	-	-	13	-	-	Quibod, M…
Muzzle Width		cm	9	9	-	-	-	3	-	-	Quibod, M…
Total Length of the Skull		cm	9	9	-	-	-	26	-	-	Quibod, M…
A1-MLL (maximum lateral length of the astragalus)	nan	mm	6	6	-	-	31	44	48	7	Wimberly,…
A2-MML (maximum medial length of the astragalus)	nan	mm	6	6	-	-	28	41	46	7	Wimberly,…
A3-MIN (minimal length of the astragalus)	nan	mm	6	6	-	-	24	35	39	6	Wimberly,…
A4-WAT (width at tarsus of the astragalus)	nan	mm	6	6	-	-	19	28	31	5	Wimberly,…
A5-WAF (width at trochlea of the astragalus)	nan	mm	6	6	-	-	17	25	28	4	Wimberly,…
C1-UL (greatest length of the calcaneus)	nan	mm	6	6	-	-	58	87	101	17	Wimberly,…
C2-TL (length of body of the calcaneus)	nan	mm	6	6	-	-	37	58	66	12	Wimberly,…
C3-MBH (midbody height of the calcaneus)	nan	mm	6	6	-	-	13	22	25	5	Wimberly,…
C4-MBTV (midbody transverse width of the calcaneus)	nan	mm	6	6	-	-	8	11	13	2	Wimberly,…
C5-FFL (fibular facet length of the calcaneus)	nan	mm	6	6	-	-	11	17	21	4	Wimberly,…
C6-CNFL (cubonavicular facet length of the calcaneus)	nan	mm	6	6	-	-	15	22	26	4	Wimberly,…
C7-HSF (height of sustentacular facet of the calcaneus)	nan	mm	6	6	-	-	13	19	21	4	Wimberly,…
C8-HST (height of sustentaculum tali of the calcaneus)	nan	mm	6	6	-	-	18	29	34	7	Wimberly,…
F1-UL (full length of the femur)	nan	mm	6	6	-	-	184	262	288	44	Wimberly,…
F2-FL (functional length of the femur)	nan	mm	6	6	-	-	185	259	282	42	Wimberly,…
F3-MCF (midshaft circumference of the femur)	nan	mm	6	6	-	-	49	72	80	13	Wimberly,…
F4-PW (proximal width of the femur)	nan	mm	6	6	-	-	41	60	67	11	Wimberly,…
F5-MLHW (mediolateral femoral head of the femur)	nan	mm	6	6	-	-	23	34	38	6	Wimberly,…
F6-APHW (anteroposterior femoral head of the femur)	nan	mm	6	6	-	-	21	30	33	5	Wimberly,…
F7-DW (distal width of the femur)	nan	mm	6	6	-	-	36	55	62	11	Wimberly,…
F8-MPL (medial condyle and patellar lip of the femur)	nan	mm	6	6	-	-	42	62	70	13	Wimberly,…
F9-LPL (lateral condyle and patellar lip of the femur)	nan	mm	6	6	-	-	39	59	65	11	Wimberly,…
H1-UL (full length of the humerus)	nan	mm	6	6	-	-	173	240	270	46	Wimberly,…
H2-FL (functional length of the humerus)	nan	mm	6	6	-	-	159	219	242	40	Wimberly,…
H3-MCF (midshaft circumference of the humerus)	nan	mm	6	6	-	-	54	81	93	18	Wimberly,…
H4-MTH (largest medial height of trochlea of the humerus)	nan	mm	6	6	-	-	19	29	34	7	Wimberly,…
H5-MidTH (smallest height of trochlea of the humerus)	nan	mm	6	6	-	-	15	23	28	5	Wimberly,…
H6-LTH (largest lateral height of trochlea of the humerus)	nan	mm	6	6	-	-	16	25	29	6	Wimberly,…
H7-PW (proximal width of the humerus)	nan	mm	6	6	-	-	39	58	67	13	Wimberly,…
H8-MLH (mediolateral humeral head width of the humerus)	nan	mm	6	6	-	-	30	45	52	10	Wimberly,…
H9-DW (distal width of the humerus)	nan	mm	6	6	-	-	31	48	57	12	Wimberly,…
MC1-UL (length of the metacarpus)	nan	mm	6	6	-	-	110	155	171	26	Wimberly,…
MC2-MAP (anteroposterior midshaft of the metacarpus)	nan	mm	6	6	-	-	11	16	18	3	Wimberly,…
MC3-MML (mediolateral midshaft of the metacarpus)	nan	mm	6	6	-	-	16	25	28	5	Wimberly,…
MC4-DW (distal width of the metacarpus)	nan	mm	6	6	-	-	25	39	44	8	Wimberly,…
MC5-PW (proximal width of the metacarpus)	nan	mm	6	6	-	-	25	38	42	7	Wimberly,…
MT1-UL (length of the metatarsus)	nan	mm	6	6	-	-	115	157	173	24	Wimberly,…
MT2-MAP (anteroposterior midshaft of the metatarsus)	nan	mm	6	6	-	-	12	17	19	3	Wimberly,…
MT3-MML (mediolateral midshaft of the metatarsus)	nan	mm	6	6	-	-	12	20	23	4	Wimberly,…
MT4-DW (distal width of the metatarsus)	nan	mm	6	6	-	-	23	36	39	7	Wimberly,…
MT5-PW (proximal width of the metatarsus)	nan	mm	6	6	-	-	21	32	36	6	Wimberly,…
R1-UL (length of the radius)	nan	mm	6	6	-	-	151	216	241	33	Wimberly,…
R2-PW (full proximal width of the radius)	nan	mm	6	6	-	-	29	45	51	8	Wimberly,…
R3-NPW (width of humeral articular surface of the radius)	nan	mm	6	6	-	-	28	44	49	8	Wimberly,…
R4-DW (distal width of the radius)	nan	mm	6	6	-	-	28	43	48	8	Wimberly,…
S1-L (full length of the scapula)	nan	mm	6	6	-	-	219	227	233	7	Wimberly,…
S2-GW (greatest cranio-caudal length of the scapula)	nan	mm	6	6	-	-	131	143	154	12	Wimberly,…
S3-SW (shortest length of scapular neck of the scapula)	nan	mm	6	6	-	-	28	31	33	3	Wimberly,…
S4-GH (anteroposterior glenoid fossa of the scapula)	nan	mm	6	6	-	-	38	40	41	2	Wimberly,…
S5-GW (mediolateral glenoid fossa of the scapula)	nan	mm	6	6	-	-	32	34	36	2	Wimberly,…
T1-L (full length of the tibia)	nan	mm	6	6	-	-	211	295	324	42	Wimberly,…
T2-MCF (midshaft circumference of the tibia)	nan	mm	6	6	-	-	46	71	78	12	Wimberly,…
T3-PW (proximal width of the tibia)	nan	mm	6	6	-	-	40	60	66	10	Wimberly,…
T4-DW (distal width of the tibia)	nan	mm	6	6	-	-	26	39	45	7	Wimberly,…
U1-UL (length of the ulna)	nan	mm	6	6	-	-	191	279	312	45	Wimberly,…
U2-OPL (olecranon process length of the ulna)	nan	mm	6	6	-	-	37	57	66	11	Wimberly,…
average ungulate dietary grass fraction (proportional)	nan	unitless	-	-	-	-	-	1	-	-	Abraham, …
femur diameter	nan	mm	-	-	-	-	-	23	-	-	Nelson, A…
humerus diameter	nan	mm	-	-	-	-	-	24	-	-	Nelson, A…

^b. other Measurements. Source Measurements which are not listed on or mapped with the Standard Traits..

Table 5. List of Data Sources for Capricornis sumatraensis.
Taxon	Reference	Match	Status
Capricornis sumatraensis	Loison,A., 1999, What factors shape sexual size dimorphism in ungulates?, Evolutionary Ecology Research	1: Exact Taxon match	Verified - Accepted
Capricornis sumatrensis	Mendoza,M., 2008, Hypsodonty in ungulates: an adaptation for grass consumption or for foraging in open habitat?, Journal of zoology	51: Name variant or misspelling of the taxon	Verified - Accepted
Capricornis sumatraensis	Grzimek,B., 2003, Grzimek's Animal Life Encyclopedia: Mammals I-V,	1: Exact Taxon match	Verified - Accepted
Capricornis sumatraensis	Quibod, M.N.R.M., Gélin, U., van Langevelde, F. and Tomlinson, K.W., 2023. Diet-specific responses of skull traits to aridity gradients in bovids and cervids. Zoological Journal of the Linnean Society, p.zlad068.	1: Exact Taxon match	Verified - Accepted
Capricornis sumatraensis	Wimberly, A.N., 2023. Predicting body mass in Ruminantia using postcranial measurements. Journal of Morphology, 284(10), p.e21636.	1: Exact Taxon match	Verified - Accepted
Capricornis sumatraensis thar	Quyoom, I., Bhat, B.A., Najar, Z.H. and Tanveer, S., 2023. Winter diet composition of Himalayan serow Capricornis sumatraensis thar in Bani Wildlife Sanctuary: implications for the conservation of Quercus semecarpifolia forest. Biologia, pp.1-9.	205: Subspecies	Verified - Accepted
Capricornis sumatraensis	Abraham, J.O., Upham, N.S., Damian-Serrano, A. and Jesmer, B.R., 2022. Evolutionary causes and consequences of ungulate migration. Nature Ecology & Evolution, 6(7), pp.998-1006.	1: Exact Taxon match	Verified - Accepted
Capricornis sumatraensis	Lundgren, E.J., Schowanek, S.D., Rowan, J., Middleton, O., Pedersen, R.Ø., Wallach, A.D., Ramp, D., Davis, M., Sandom, C.J. and Svenning, J.C., 2021. Functional traits of the world’s late Quaternary large-bodied avian and mammalian herbivores. Scientific data, 8(1), p.17.	1: Exact Taxon match	Verified - Accepted
Capricornis sumatraensis	Nelson, A., Engelman, R.K. and Croft, D.A., 2023. How to weigh a fossil mammal? South American notoungulates as a case study for estimating body mass in extinct clades. Journal of Mammalian Evolution, 30(3), pp.773-809.	1: Exact Taxon match	Verified - Accepted
Capricornis sumatraensis	Freudenthal, M. and Martín-Suárez, E., 2013. Estimating body mass of fossil rodents. Scripta Geologica, 145, pp.1-513.	1: Exact Taxon match	Verified - Accepted
Capricornis sumatraensis	Beaudrot, L., Acevedo, M.A., Gorczynski, D. and Harris, N.C., 2024. Geographic differences in body size distributions underlie food web connectance of tropical forest mammals. Scientific Reports, 14(1), pp.1-10.	1: Exact Taxon match	Verified - Accepted
Capricornis sumatraensis	Rowan, J., Beaudrot, L., Franklin, J., Reed, K.E., Smail, I.E., Zamora, A. and Kamilar, J.M., 2020. Geographically divergent evolutionary and ecological legacies shape mammal biodiversity in the global tropics and subtropics. Proceedings of the National Academy of Sciences, 117(3), pp.1559-1565.	1: Exact Taxon match	Verified - Accepted

Citation for this page: [The MammalBase community 2026. / CC BY 4.0. http://doi.org/10.5281/zenodo.7462864 Capricornis sumatraensis , Accessed 4th April 2026 at https://mammalbase.net/me/16081]