MammalBase - Accepted taxon: Tragelaphus scriptus

Accepted taxon: Tragelaphus scriptus
Taxon:	Tragelaphus scriptus
Rank:	Species
Higher Classification:	ARTIODACTYLA \| Bovidae \| Bovinae \| Tragelaphus
Reference:	Wilson, Don E., and DeeAnn M. Reeder, eds. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), JHU Press, 2005.

Fig.1 - Ternary plot of the proximate analysis for the Tragelaphus scriptus diet.
CF (Crude Fibre) 24, ASH (Ash contents) 10, NFE (Nitrogen-Free-Extract) 51, CP (Crude Protein) 11, EE (Ether Extract) 4. See details for the calculations in Lintulaakso, K., Tatti, N. and Žliobaitė, I., 2022. Quantifying mammalian diets. Mammalian Biology, pp.1-15. [DOI]

Table 1. List of Diet Items for Tragelaphus scriptus.
TSN Complete name	Part	Ratio ^a.	Reference(s)
Magnoliopsida	SHOOT	21.33	Damuth, J…
Magnoliopsida	LEAF	16.67	Grzimek,B…
Magnoliopsida	STEM	12.00	Grzimek,B…
Poaceae	STEM	8.00	Damuth, J…
Maytenus	LEAF	5.13	Bayih, W.…
Maytenus	FRUIT	2.70	Bayih, W.…
Maytenus	SHOOT	2.43	Bayih, W.…
Cyperus	LEAF	2.33	Bayih, W.…
Rosa abyssinica	LEAF	2.12	Bayih, W.…
Magnoliopsida	FLOWER	2.00	Grzimek,B…
Carissa	FRUIT	1.90	Bayih, W.…
Urera	LEAF	1.80	Bayih, W.…
Carissa	LEAF	1.80	Bayih, W.…
Cynodon dactylon	LEAF	1.69	Bayih, W.…
Clematis	LEAF	1.64	Bayih, W.…
Cynodon dactylon	STEM	1.59	Bayih, W.…
Rosacea	LEAF	1.53	Bayih, W.…
Veronica	LEAF	1.27	Bayih, W.…
Albizia gummifera	LEAF	1.27	Bayih, W.…
Proteaceae	LEAF	1.22	Bayih, W.…
Plantae	SHOOT	1.22	Bayih, W.…
Polystachya	LEAF	1.11	Bayih, W.…
Cynodon nlemfuensis	LEAF	1.06	Bayih, W.…
Dombeya	LEAF	1.01	Bayih, W.…
Polystachya	SHOOT	1.01	Bayih, W.…
Cynodon nlemfuensis	STEM	0.95	Bayih, W.…
Sesbania sesban	LEAF	0.74	Bayih, W.…
Echinops	LEAF	0.69	Bayih, W.…
Acacia abyssinica	LEAF	0.69	Bayih, W.…
Acacia abyssinica	SHOOT	0.58	Bayih, W.…
Eucalyptus camaldulensis	LEAF	0.53	Bayih, W.…

^a. Standardized values. See details for the calculations in Lintulaakso, K., Tatti, N. and Žliobaitė, I., 2022. Quantifying mammalian diets. Mammalian Biology, pp.1-15. [DOI]

Table 2. List of Standard Traits for Tragelaphus scriptus.
Trait	Selected Value	All Value(s)	Reference(s)
Dietary category: Eisenberg 1981	Herbivore-Browser	Herbivore-Browser	– Abraham, J.O., Upham, N.S., Damian-Serrano, A. and Jesmer, B.R., 2022. Evolutionary causes and consequences of ungulate migration. Nature Ecology & Evolution, 6(7), pp.998-1006., I. J. Gordon, A. W. Illius, 1988, 'Incisor Arcade Structure and Diet Selection in Ruminants', Functional Ecology, vol. 2, no. 1, p. 15, Loison,A., 1999, What factors shape sexual size dimorphism in ungulates?, Evolutionary Ecology Research, M. Clauss, R. R. Hofmann, W. J. Streich, J. Fickel, J. Hummel, 2009, 'Convergence in the macroscopic anatomy of the reticulum in wild ruminant species of different feeding types and a new resulting hypothesis on reticular function', Journal of Zoology, vol. 281, no. 1, pp. 26-38, M. Mendoza, P. Palmqvist, 2008, 'Hypsodonty in ungulates: an adaptation for grass consumption or for foraging in open habitat?', Journal of Zoology, vol. 274, no. 2, pp. 134-142, Quibod, M., Gélin, U., van Langevelde, F., Tomlinson, K. 2023. Diet-specific responses of skull traits to aridity gradients in bovids and cervids. Zoological Journal of the Linnean Society. None(None), pp.None. Available at: 10.1093/zoolinnean/zlad068.
Dietary category: Miljutin 2009	Herbivore	Herbivore	– Abraham, J.O., Upham, N.S., Damian-Serrano, A. and Jesmer, B.R., 2022. Evolutionary causes and consequences of ungulate migration. Nature Ecology & Evolution, 6(7), pp.998-1006., I. J. Gordon, A. W. Illius, 1988, 'Incisor Arcade Structure and Diet Selection in Ruminants', Functional Ecology, vol. 2, no. 1, p. 15, Loison,A., 1999, What factors shape sexual size dimorphism in ungulates?, Evolutionary Ecology Research, M. Clauss, R. R. Hofmann, W. J. Streich, J. Fickel, J. Hummel, 2009, 'Convergence in the macroscopic anatomy of the reticulum in wild ruminant species of different feeding types and a new resulting hypothesis on reticular function', Journal of Zoology, vol. 281, no. 1, pp. 26-38, M. Mendoza, P. Palmqvist, 2008, 'Hypsodonty in ungulates: an adaptation for grass consumption or for foraging in open habitat?', Journal of Zoology, vol. 274, no. 2, pp. 134-142, Quibod, M., Gélin, U., van Langevelde, F., Tomlinson, K. 2023. Diet-specific responses of skull traits to aridity gradients in bovids and cervids. Zoological Journal of the Linnean Society. None(None), pp.None. Available at: 10.1093/zoolinnean/zlad068.

Table 3. List of Standard Measurements for Tragelaphus scriptus.
Measurement	Location	Unit	n Total	n Unknown	n Female	n Male	Min	Mean	Max	SD	Reference(s)
Body Mass (BM)	nan	g	-	-	-	-	-	48,340	-	-	Wimberly,…
Body Mass (BM)	nan	g	-	-	-	-	21,000	-	80,000	-	Freudenth…

Table 4. List of other Measurements ^b. for Tragelaphus scriptus.
Measurement	Location	Unit	n Total	n Unknown	n Female	n Male	Min	Mean	Max	SD	Reference(s)
Length of the Masseteric Fossa		cm	12	12	-	-	-	11	-	-	Quibod, M…
Muzzle Width		cm	12	12	-	-	-	3	-	-	Quibod, M…
Total Length of the Skull		cm	12	12	-	-	-	21	-	-	Quibod, M…
A1-MLL (maximum lateral length of the astragalus)	nan	mm	6	6	-	-	33	37	41	3	Wimberly,…
A2-MML (maximum medial length of the astragalus)	nan	mm	6	6	-	-	31	36	39	3	Wimberly,…
A3-MIN (minimal length of the astragalus)	nan	mm	6	6	-	-	26	30	33	3	Wimberly,…
A4-WAT (width at tarsus of the astragalus)	nan	mm	6	6	-	-	19	22	24	2	Wimberly,…
A5-WAF (width at trochlea of the astragalus)	nan	mm	6	6	-	-	18	21	23	2	Wimberly,…
C1-UL (greatest length of the calcaneus)	nan	mm	6	6	-	-	69	77	87	7	Wimberly,…
C2-TL (length of body of the calcaneus)	nan	mm	6	6	-	-	45	51	61	6	Wimberly,…
C3-MBH (midbody height of the calcaneus)	nan	mm	6	6	-	-	18	21	26	3	Wimberly,…
C4-MBTV (midbody transverse width of the calcaneus)	nan	mm	6	6	-	-	11	12	14	1	Wimberly,…
C5-FFL (fibular facet length of the calcaneus)	nan	mm	6	6	-	-	14	15	17	1	Wimberly,…
C6-CNFL (cubonavicular facet length of the calcaneus)	nan	mm	6	6	-	-	17	19	20	1	Wimberly,…
C7-HSF (height of sustentacular facet of the calcaneus)	nan	mm	6	6	-	-	14	15	16	1	Wimberly,…
C8-HST (height of sustentaculum tali of the calcaneus)	nan	mm	6	6	-	-	21	25	28	3	Wimberly,…
F1-UL (full length of the femur)	nan	mm	6	6	-	-	222	242	271	21	Wimberly,…
F2-FL (functional length of the femur)	nan	mm	6	6	-	-	214	232	257	18	Wimberly,…
F3-MCF (midshaft circumference of the femur)	nan	mm	6	6	-	-	60	69	79	9	Wimberly,…
F4-PW (proximal width of the femur)	nan	mm	6	6	-	-	45	53	59	6	Wimberly,…
F5-MLHW (mediolateral femoral head of the femur)	nan	mm	6	6	-	-	29	33	36	3	Wimberly,…
F6-APHW (anteroposterior femoral head of the femur)	nan	mm	6	6	-	-	21	24	28	3	Wimberly,…
F7-DW (distal width of the femur)	nan	mm	6	6	-	-	43	46	50	3	Wimberly,…
F8-MPL (medial condyle and patellar lip of the femur)	nan	mm	6	6	-	-	49	58	66	7	Wimberly,…
F9-LPL (lateral condyle and patellar lip of the femur)	nan	mm	6	6	-	-	45	53	61	7	Wimberly,…
H1-UL (full length of the humerus)	nan	mm	6	6	-	-	171	188	212	15	Wimberly,…
H2-FL (functional length of the humerus)	nan	mm	6	6	-	-	157	170	191	13	Wimberly,…
H3-MCF (midshaft circumference of the humerus)	nan	mm	6	6	-	-	57	64	74	8	Wimberly,…
H4-MTH (largest medial height of trochlea of the humerus)	nan	mm	6	6	-	-	22	24	26	2	Wimberly,…
H5-MidTH (smallest height of trochlea of the humerus)	nan	mm	6	6	-	-	16	18	20	2	Wimberly,…
H6-LTH (largest lateral height of trochlea of the humerus)	nan	mm	6	6	-	-	18	20	23	2	Wimberly,…
H7-PW (proximal width of the humerus)	nan	mm	6	6	-	-	37	42	49	6	Wimberly,…
H8-MLH (mediolateral humeral head width of the humerus)	nan	mm	6	6	-	-	29	34	41	4	Wimberly,…
H9-DW (distal width of the humerus)	nan	mm	6	6	-	-	33	36	39	3	Wimberly,…
MC1-UL (length of the metacarpus)	nan	mm	6	6	-	-	140	168	191	19	Wimberly,…
MC2-MAP (anteroposterior midshaft of the metacarpus)	nan	mm	6	6	-	-	12	13	15	1	Wimberly,…
MC3-MML (mediolateral midshaft of the metacarpus)	nan	mm	6	6	-	-	13	15	18	2	Wimberly,…
MC4-DW (distal width of the metacarpus)	nan	mm	6	6	-	-	21	24	27	2	Wimberly,…
MC5-PW (proximal width of the metacarpus)	nan	mm	6	6	-	-	23	24	28	2	Wimberly,…
MT1-UL (length of the metatarsus)	nan	mm	6	6	-	-	153	180	201	18	Wimberly,…
MT2-MAP (anteroposterior midshaft of the metatarsus)	nan	mm	6	6	-	-	15	18	21	3	Wimberly,…
MT3-MML (mediolateral midshaft of the metatarsus)	nan	mm	6	6	-	-	13	16	18	2	Wimberly,…
MT4-DW (distal width of the metatarsus)	nan	mm	6	6	-	-	23	27	31	3	Wimberly,…
MT5-PW (proximal width of the metatarsus)	nan	mm	6	6	-	-	22	24	26	2	Wimberly,…
R1-UL (length of the radius)	nan	mm	6	6	-	-	160	178	205	17	Wimberly,…
R2-PW (full proximal width of the radius)	nan	mm	6	6	-	-	31	33	38	3	Wimberly,…
R3-NPW (width of humeral articular surface of the radius)	nan	mm	6	6	-	-	29	31	34	2	Wimberly,…
R4-DW (distal width of the radius)	nan	mm	6	6	-	-	28	29	33	2	Wimberly,…
S1-L (full length of the scapula)	nan	mm	6	6	-	-	147	166	197	27	Wimberly,…
S2-GW (greatest cranio-caudal length of the scapula)	nan	mm	6	6	-	-	85	97	114	15	Wimberly,…
S3-SW (shortest length of scapular neck of the scapula)	nan	mm	6	6	-	-	16	19	23	4	Wimberly,…
S4-GH (anteroposterior glenoid fossa of the scapula)	nan	mm	6	6	-	-	24	27	31	4	Wimberly,…
S5-GW (mediolateral glenoid fossa of the scapula)	nan	mm	6	6	-	-	21	25	29	4	Wimberly,…
T1-L (full length of the tibia)	nan	mm	6	6	-	-	244	269	286	22	Wimberly,…
T2-MCF (midshaft circumference of the tibia)	nan	mm	6	6	-	-	62	71	77	8	Wimberly,…
T3-PW (proximal width of the tibia)	nan	mm	6	6	-	-	48	53	57	5	Wimberly,…
T4-DW (distal width of the tibia)	nan	mm	6	6	-	-	31	34	37	3	Wimberly,…
U1-UL (length of the ulna)	nan	mm	6	6	-	-	202	222	258	22	Wimberly,…
U2-OPL (olecranon process length of the ulna)	nan	mm	6	6	-	-	37	40	49	5	Wimberly,…
average ungulate dietary grass fraction (proportional)	nan	unitless	-	-	-	-	-	0	-	-	Abraham, …
femur circumference	nan	mm	-	-	-	-	-	56	-	-	Nelson, A…
femur diameter	nan	mm	-	-	-	-	-	21	-	-	Nelson, A…
humerus circumference	nan	mm	-	-	-	-	-	49	-	-	Nelson, A…
humerus diameter	nan	mm	-	-	-	-	-	17	-	-	Nelson, A…
neck length	nan	mm	-	-	-	-	-	348	-	-	Nelson, A…

^b. other Measurements. Source Measurements which are not listed on or mapped with the Standard Traits..

Table 5. List of Data Sources for Tragelaphus scriptus.
Taxon	Reference	Match	Status
Tragelaphus scriptus	Clauss,M., 2010, Convergence in the macroscopic anatomy of the reticulum in wild ruminant species of different feeding types and a new resulting hypothesis on reticular function, Journal of zoology	1: Exact Taxon match	Verified - Accepted
Tragelaphus scriptus	Gordon,IJ, 1988, Incisor arcade structure and diet selection in ruminants, Functional Ecology	1: Exact Taxon match	Verified - Accepted
Tragelaphus scriptus	Loison,A., 1999, What factors shape sexual size dimorphism in ungulates?, Evolutionary Ecology Research	1: Exact Taxon match	Verified - Accepted
Tragelaphus scriptus	Mendoza,M., 2008, Hypsodonty in ungulates: an adaptation for grass consumption or for foraging in open habitat?, Journal of zoology	1: Exact Taxon match	Verified - Accepted
Tragelaphus scriptus	Damuth, J. 2010. ABSOLUT 6 a mammalian database compiled by the national center for ecological analysis and synthesis (NCEAS ) workshop on mammalian communities.	1: Exact Taxon match	Verified - Accepted
Tragelaphus scriptus	Nowak,R. M., 1991, Walker's mammals of the world,	1: Exact Taxon match	Verified - Accepted
Tragelaphus scriptus	Grzimek,B., 2003, Grzimek's Animal Life Encyclopedia: Mammals I-V,	1: Exact Taxon match	Verified - Accepted
Tragelaphus scriptus	Nowak,R. M., 1991, Walker's mammals of the world,	1: Exact Taxon match	Verified - Accepted
Tragelaphus scriptus decula	Bayih, W. and Yihune, M., 2018. Population status, feeding ecology and activity pattern of common bushbuck (Tragelaphus Scriptus decula) in Sekele Mariam Forest, West Gojjam, Ethiopia. Journal of Ecology and The Natural Environment, 10(5), pp.69-79.	205: Subspecies	Verified - Accepted
Tragelaphus scriptus	Quibod, M.N.R.M., Gélin, U., van Langevelde, F. and Tomlinson, K.W., 2023. Diet-specific responses of skull traits to aridity gradients in bovids and cervids. Zoological Journal of the Linnean Society, p.zlad068.	1: Exact Taxon match	Verified - Accepted
Tragelaphus scriptus	DeMers, A.C., 2023. Functional Significance of the Mandible, Tooth Roots, and Tooth Crowns, and their Implications for Fossil Dietary Inference (Doctoral dissertation, The Ohio State University).	1: Exact Taxon match	Verified - Accepted
Tragelaphus scriptus	Wimberly, A.N., 2023. Predicting body mass in Ruminantia using postcranial measurements. Journal of Morphology, 284(10), p.e21636.	1: Exact Taxon match	Verified - Accepted
Tragelaphus scriptus	Abraham, J.O., Upham, N.S., Damian-Serrano, A. and Jesmer, B.R., 2022. Evolutionary causes and consequences of ungulate migration. Nature Ecology & Evolution, 6(7), pp.998-1006.	1: Exact Taxon match	Verified - Accepted
Tragelaphus scriptus	Lundgren, E.J., Schowanek, S.D., Rowan, J., Middleton, O., Pedersen, R.Ø., Wallach, A.D., Ramp, D., Davis, M., Sandom, C.J. and Svenning, J.C., 2021. Functional traits of the world’s late Quaternary large-bodied avian and mammalian herbivores. Scientific data, 8(1), p.17.	1: Exact Taxon match	Verified - Accepted
Tragelaphus scriptus	Nelson, A., Engelman, R.K. and Croft, D.A., 2023. How to weigh a fossil mammal? South American notoungulates as a case study for estimating body mass in extinct clades. Journal of Mammalian Evolution, 30(3), pp.773-809.	1: Exact Taxon match	Verified - Accepted
Tragelaphus scriptus	Freudenthal, M. and Martín-Suárez, E., 2013. Estimating body mass of fossil rodents. Scripta Geologica, 145, pp.1-513.	1: Exact Taxon match	Verified - Accepted
Tragelaphus scriptus	Beaudrot, L., Acevedo, M.A., Gorczynski, D. and Harris, N.C., 2024. Geographic differences in body size distributions underlie food web connectance of tropical forest mammals. Scientific Reports, 14(1), pp.1-10.	1: Exact Taxon match	Verified - Accepted
Tragelaphus scriptus	Rowan, J., Beaudrot, L., Franklin, J., Reed, K.E., Smail, I.E., Zamora, A. and Kamilar, J.M., 2020. Geographically divergent evolutionary and ecological legacies shape mammal biodiversity in the global tropics and subtropics. Proceedings of the National Academy of Sciences, 117(3), pp.1559-1565.	1: Exact Taxon match	Verified - Accepted

Citation for this page: [The MammalBase community 2026. / CC BY 4.0. http://doi.org/10.5281/zenodo.7462864 Tragelaphus scriptus , Accessed 15th April 2026 at https://mammalbase.net/me/16023]