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MammalBase - Accepted taxon: Raphicerus campestris

Accepted taxon: Raphicerus campestris


Accepted taxon: Raphicerus campestris
Taxon: Raphicerus campestris
Rank: Species
Higher Classification: ARTIODACTYLA | Bovidae | Antilopinae | Raphicerus
Reference: Wilson, Don E., and DeeAnn M. Reeder, eds. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), JHU Press, 2005.

Fig.1 - Ternary plot of the proximate analysis for the Raphicerus campestris diet.
CF (Crude Fibre) 19, ASH (Ash contents) 7, NFE (Nitrogen-Free-Extract) 57, CP (Crude Protein) 11, EE (Ether Extract) 4. See details for the calculations in Lintulaakso, K., Tatti, N. and Žliobaitė, I., 2022. Quantifying mammalian diets. Mammalian Biology, pp.1-15. [DOI]


Table 1. List of Diet Items for Raphicerus campestris.
TSN Complete name Part Ratio a. Reference(s)
Magnoliopsida LEAF 28.57 Damuth, J…
Magnoliopsida ROOT 17.62 Damuth, J…
Magnoliopsida FRUIT 14.76 Damuth, J…
Poaceae STEM 13.33 Damuth, J…
Magnoliopsida SHOOT 9.52 Grzimek,B…
Magnoliopsida FLOWER 7.14 Grzimek,B…
Magnoliopsida SEED 6.67 Damuth, J…
0 MINERAL 2.38 Grzimek,B…

a. Standardized values. See details for the calculations in Lintulaakso, K., Tatti, N. and Žliobaitė, I., 2022. Quantifying mammalian diets. Mammalian Biology, pp.1-15. [DOI]


Table 2. List of Standard Traits for Raphicerus campestris.
Trait Selected Value All Value(s) Reference(s)
Dietary category: Eisenberg 1981 Herbivore-Mixed feeder Herbivore-Browser, Herbivore-Mixed feeder Abraham, J.O., Upham, N.S., Damian-Serrano, A. and Jesmer, B.R., 2022. Evolutionary causes and consequences of ungulate migration. Nature Ecology & Evolution, 6(7), pp.998-1006., I. J. Gordon, A. W. Illius, 1988, 'Incisor Arcade Structure and Diet Selection in Ruminants', Functional Ecology, vol. 2, no. 1, p. 15, Loison,A., 1999, What factors shape sexual size dimorphism in ungulates?, Evolutionary Ecology Research, M. Clauss, R. R. Hofmann, W. J. Streich, J. Fickel, J. Hummel, 2009, 'Convergence in the macroscopic anatomy of the reticulum in wild ruminant species of different feeding types and a new resulting hypothesis on reticular function', Journal of Zoology, vol. 281, no. 1, pp. 26-38, M. Mendoza, P. Palmqvist, 2008, 'Hypsodonty in ungulates: an adaptation for grass consumption or for foraging in open habitat?', Journal of Zoology, vol. 274, no. 2, pp. 134-142, Quibod, M., Gélin, U., van Langevelde, F., Tomlinson, K. 2023. Diet-specific responses of skull traits to aridity gradients in bovids and cervids. Zoological Journal of the Linnean Society. None(None), pp.None. Available at: 10.1093/zoolinnean/zlad068.
Dietary category: Miljutin 2009 Herbivore Herbivore Abraham, J.O., Upham, N.S., Damian-Serrano, A. and Jesmer, B.R., 2022. Evolutionary causes and consequences of ungulate migration. Nature Ecology & Evolution, 6(7), pp.998-1006., I. J. Gordon, A. W. Illius, 1988, 'Incisor Arcade Structure and Diet Selection in Ruminants', Functional Ecology, vol. 2, no. 1, p. 15, Loison,A., 1999, What factors shape sexual size dimorphism in ungulates?, Evolutionary Ecology Research, M. Clauss, R. R. Hofmann, W. J. Streich, J. Fickel, J. Hummel, 2009, 'Convergence in the macroscopic anatomy of the reticulum in wild ruminant species of different feeding types and a new resulting hypothesis on reticular function', Journal of Zoology, vol. 281, no. 1, pp. 26-38, M. Mendoza, P. Palmqvist, 2008, 'Hypsodonty in ungulates: an adaptation for grass consumption or for foraging in open habitat?', Journal of Zoology, vol. 274, no. 2, pp. 134-142, Quibod, M., Gélin, U., van Langevelde, F., Tomlinson, K. 2023. Diet-specific responses of skull traits to aridity gradients in bovids and cervids. Zoological Journal of the Linnean Society. None(None), pp.None. Available at: 10.1093/zoolinnean/zlad068.

Table 3. List of Standard Measurements for Raphicerus campestris.
Measurement Location Unit n Total n Unknown n Female n Male Min Mean Max SD Reference(s)
Body Mass (BM) nan g 10 10 - - - 10,545.45 - - Engelman,…
Body Mass (BM) nan g - - - - - 11,290 - - Wimberly,…
Body Mass (BM) nan g - - - - 7,000 - 16,000 - Freudenth…
Head-Body Length (HBL) nan mm - - - - - 795 - - Engelman,…
Occipital condyle width (OCW) nan mm 10 10 - - - 27.28 - - Engelman,…

Table 4. List of other Measurements b. for Raphicerus campestris.
Measurement Location Unit n Total n Unknown n Female n Male Min Mean Max SD Reference(s)
Length of the Masseteric Fossa cm 8 8 - - - 7 - - Quibod, M…
Muzzle Width cm 8 8 - - - 1 - - Quibod, M…
Total Length of the Skull cm 8 8 - - - 14 - - Quibod, M…
brainmass nan g - - - - - 51 - - Engelman,…
skull_length nan mm - - - - - 128 - - Engelman,…
A1-MLL (maximum lateral length of the astragalus) nan mm 7 7 - - 22 23 24 1 Wimberly,…
A2-MML (maximum medial length of the astragalus) nan mm 7 7 - - 21 22 23 1 Wimberly,…
A3-MIN (minimal length of the astragalus) nan mm 7 7 - - 18 19 19 1 Wimberly,…
A4-WAT (width at tarsus of the astragalus) nan mm 7 7 - - 13 14 15 0 Wimberly,…
A5-WAF (width at trochlea of the astragalus) nan mm 7 7 - - 12 13 14 1 Wimberly,…
C1-UL (greatest length of the calcaneus) nan mm 7 7 - - 45 48 49 2 Wimberly,…
C2-TL (length of body of the calcaneus) nan mm 7 7 - - 29 32 34 2 Wimberly,…
C3-MBH (midbody height of the calcaneus) nan mm 7 7 - - 12 13 13 1 Wimberly,…
C4-MBTV (midbody transverse width of the calcaneus) nan mm 7 7 - - 6 7 7 0 Wimberly,…
C5-FFL (fibular facet length of the calcaneus) nan mm 7 7 - - 9 10 11 1 Wimberly,…
C6-CNFL (cubonavicular facet length of the calcaneus) nan mm 7 7 - - 11 12 13 1 Wimberly,…
C7-HSF (height of sustentacular facet of the calcaneus) nan mm 7 7 - - 9 9 10 1 Wimberly,…
C8-HST (height of sustentaculum tali of the calcaneus) nan mm 7 7 - - 13 14 16 1 Wimberly,…
F1-UL (full length of the femur) nan mm 7 7 - - 141 148 155 6 Wimberly,…
F2-FL (functional length of the femur) nan mm 7 7 - - 134 142 148 6 Wimberly,…
F3-MCF (midshaft circumference of the femur) nan mm 7 7 - - 37 40 43 3 Wimberly,…
F4-PW (proximal width of the femur) nan mm 7 7 - - 33 36 38 2 Wimberly,…
F5-MLHW (mediolateral femoral head of the femur) nan mm 7 7 - - 20 20 21 1 Wimberly,…
F6-APHW (anteroposterior femoral head of the femur) nan mm 7 7 - - 15 15 16 1 Wimberly,…
F7-DW (distal width of the femur) nan mm 7 7 - - 28 29 30 1 Wimberly,…
F8-MPL (medial condyle and patellar lip of the femur) nan mm 7 7 - - 38 38 40 1 Wimberly,…
F9-LPL (lateral condyle and patellar lip of the femur) nan mm 7 7 - - 33 34 35 1 Wimberly,…
H1-UL (full length of the humerus) nan mm 7 7 - - 101 105 110 3 Wimberly,…
H2-FL (functional length of the humerus) nan mm 7 7 - - 95 98 103 3 Wimberly,…
H3-MCF (midshaft circumference of the humerus) nan mm 7 7 - - 30 33 35 2 Wimberly,…
H4-MTH (largest medial height of trochlea of the humerus) nan mm 7 7 - - 14 14 14 0 Wimberly,…
H5-MidTH (smallest height of trochlea of the humerus) nan mm 7 7 - - 10 10 11 0 Wimberly,…
H6-LTH (largest lateral height of trochlea of the humerus) nan mm 7 7 - - 11 12 12 0 Wimberly,…
H7-PW (proximal width of the humerus) nan mm 7 7 - - 23 25 26 1 Wimberly,…
H8-MLH (mediolateral humeral head width of the humerus) nan mm 7 7 - - 18 19 21 1 Wimberly,…
H9-DW (distal width of the humerus) nan mm 7 7 - - 19 20 21 1 Wimberly,…
MC1-UL (length of the metacarpus) nan mm 7 7 - - 120 125 132 4 Wimberly,…
MC2-MAP (anteroposterior midshaft of the metacarpus) nan mm 7 7 - - 7 7 8 0 Wimberly,…
MC3-MML (mediolateral midshaft of the metacarpus) nan mm 7 7 - - 8 9 10 1 Wimberly,…
MC4-DW (distal width of the metacarpus) nan mm 7 7 - - 14 15 15 0 Wimberly,…
MC5-PW (proximal width of the metacarpus) nan mm 7 7 - - 15 16 17 1 Wimberly,…
MT1-UL (length of the metatarsus) nan mm 7 7 - - 126 131 136 3 Wimberly,…
MT2-MAP (anteroposterior midshaft of the metatarsus) nan mm 7 7 - - 10 11 12 1 Wimberly,…
MT3-MML (mediolateral midshaft of the metatarsus) nan mm 7 7 - - 8 9 10 1 Wimberly,…
MT4-DW (distal width of the metatarsus) nan mm 7 7 - - 17 17 18 0 Wimberly,…
MT5-PW (proximal width of the metatarsus) nan mm 7 7 - - 16 16 17 1 Wimberly,…
R1-UL (length of the radius) nan mm 7 7 - - 114 118 125 4 Wimberly,…
R2-PW (full proximal width of the radius) nan mm 7 7 - - 18 18 20 1 Wimberly,…
R3-NPW (width of humeral articular surface of the radius) nan mm 7 7 - - 17 17 18 1 Wimberly,…
R4-DW (distal width of the radius) nan mm 7 7 - - 16 17 18 1 Wimberly,…
S1-L (full length of the scapula) nan mm 7 7 - - 87 90 94 5 Wimberly,…
S2-GW (greatest cranio-caudal length of the scapula) nan mm 7 7 - - 56 57 58 1 Wimberly,…
S3-SW (shortest length of scapular neck of the scapula) nan mm 7 7 - - 9 9 10 1 Wimberly,…
S4-GH (anteroposterior glenoid fossa of the scapula) nan mm 7 7 - - 15 16 16 1 Wimberly,…
S5-GW (mediolateral glenoid fossa of the scapula) nan mm 7 7 - - 12 13 14 1 Wimberly,…
T1-L (full length of the tibia) nan mm 7 7 - - 171 177 187 6 Wimberly,…
T2-MCF (midshaft circumference of the tibia) nan mm 7 7 - - 38 41 45 2 Wimberly,…
T3-PW (proximal width of the tibia) nan mm 7 7 - - 29 31 32 1 Wimberly,…
T4-DW (distal width of the tibia) nan mm 7 7 - - 19 20 21 1 Wimberly,…
U1-UL (length of the ulna) nan mm 7 7 - - 137 143 152 6 Wimberly,…
U2-OPL (olecranon process length of the ulna) nan mm 7 7 - - 22 24 27 2 Wimberly,…
average ungulate dietary grass fraction (proportional) nan unitless - - - - - 0 - - Abraham, …
BMR (kJ/h) nan kJ/h - - - - - 66 - - McNab, B.…
femur diameter nan mm - - - - - 12 - - Nelson, A…
humerus diameter nan mm - - - - - 9 - - Nelson, A…
neck length nan mm - - - - - 146 - - Nelson, A…

b. other Measurements. Source Measurements which are not listed on or mapped with the Standard Traits..


Table 5. List of Data Sources for Raphicerus campestris.
Taxon Reference Match Status
Raphicerus campestris Clauss,M., 2010, Convergence in the macroscopic anatomy of the reticulum in wild ruminant species of different feeding types and a new resulting hypothesis on reticular function, Journal of zoology 1: Exact Taxon match Verified - Accepted
Raphicerus campestris Gordon,IJ, 1988, Incisor arcade structure and diet selection in ruminants, Functional Ecology 1: Exact Taxon match Verified - Accepted
Raphicerus campestris Loison,A., 1999, What factors shape sexual size dimorphism in ungulates?, Evolutionary Ecology Research 1: Exact Taxon match Verified - Accepted
Raphicerus campestris Mendoza,M., 2008, Hypsodonty in ungulates: an adaptation for grass consumption or for foraging in open habitat?, Journal of zoology 1: Exact Taxon match Verified - Accepted
Raphicerus campestris Damuth, J. 2010. ABSOLUT 6 a mammalian database compiled by the national center for ecological analysis and synthesis (NCEAS ) workshop on mammalian communities. 1: Exact Taxon match Verified - Accepted
Raphicerus campestris Grzimek,B., 2003, Grzimek's Animal Life Encyclopedia: Mammals I-V, 1: Exact Taxon match Verified - Accepted
Raphicerus campestris Quibod, M.N.R.M., Gélin, U., van Langevelde, F. and Tomlinson, K.W., 2023. Diet-specific responses of skull traits to aridity gradients in bovids and cervids. Zoological Journal of the Linnean Society, p.zlad068. 1: Exact Taxon match Verified - Accepted
Raphicerus campestris Engelman, R.K., 2022. Occipital condyle width (OCW) is a highly accurate predictor of body mass in therian mammals. BMC biology, 20(1), p.37. 1: Exact Taxon match Verified - Accepted
Raphicerus campestris Wimberly, A.N., 2023. Predicting body mass in Ruminantia using postcranial measurements. Journal of Morphology, 284(10), p.e21636. 1: Exact Taxon match Verified - Accepted
Raphicerus campestris Abraham, J.O., Upham, N.S., Damian-Serrano, A. and Jesmer, B.R., 2022. Evolutionary causes and consequences of ungulate migration. Nature Ecology & Evolution, 6(7), pp.998-1006. 1: Exact Taxon match Verified - Accepted
Raphicerus campestris Lundgren, E.J., Schowanek, S.D., Rowan, J., Middleton, O., Pedersen, R.Ø., Wallach, A.D., Ramp, D., Davis, M., Sandom, C.J. and Svenning, J.C., 2021. Functional traits of the world’s late Quaternary large-bodied avian and mammalian herbivores. Scientific data, 8(1), p.17. 1: Exact Taxon match Verified - Accepted
Raphicerus campestris McNab, B.K., 2008. An analysis of the factors that influence the level and scaling of mammalian BMR. Comparative Biochemistry and Physiology Part A: Molecular & Integrative Physiology, 151(1), pp.5-28. 1: Exact Taxon match Verified - Accepted
Raphicerus campestris Nelson, A., Engelman, R.K. and Croft, D.A., 2023. How to weigh a fossil mammal? South American notoungulates as a case study for estimating body mass in extinct clades. Journal of Mammalian Evolution, 30(3), pp.773-809. 1: Exact Taxon match Verified - Accepted
Raphicerus campestris Freudenthal, M. and Martín-Suárez, E., 2013. Estimating body mass of fossil rodents. Scripta Geologica, 145, pp.1-513. 1: Exact Taxon match Verified - Accepted
Raphicerus campestris Rowan, J., Beaudrot, L., Franklin, J., Reed, K.E., Smail, I.E., Zamora, A. and Kamilar, J.M., 2020. Geographically divergent evolutionary and ecological legacies shape mammal biodiversity in the global tropics and subtropics. Proceedings of the National Academy of Sciences, 117(3), pp.1559-1565. 1: Exact Taxon match Verified - Accepted

Citation for this page: [The MammalBase community 2026. / CC BY 4.0. http://doi.org/10.5281/zenodo.7462864 Raphicerus campestris , Accessed 15th April 2026 at https://mammalbase.net/me/15958]