MammalBase - Accepted taxon: Helarctos malayanus

Accepted taxon: Helarctos malayanus
Taxon:	Helarctos malayanus
Rank:	Species
Higher Classification:	CARNIVORA \| CANIFORMIA \| Ursidae \| Helarctos
Reference:	Wilson, Don E., and DeeAnn M. Reeder, eds. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), JHU Press, 2005.

Fig.1 - Ternary plot of the proximate analysis for the Helarctos malayanus diet.
CF (Crude Fibre) 8, ASH (Ash contents) 6, NFE (Nitrogen-Free-Extract) 45, CP (Crude Protein) 29, EE (Ether Extract) 13. See details for the calculations in Lintulaakso, K., Tatti, N. and Žliobaitė, I., 2022. Quantifying mammalian diets. Mammalian Biology, pp.1-15. [DOI]

Table 1. List of Diet Items for Helarctos malayanus.
TSN Complete name	Part	Ratio ^a.	Reference(s)
Cocos nucifera	FRUIT	13.07	Fitzgeral…
Magnoliopsida	EXUDATES	10.42	Fitzgeral…
Termitidae	WHOLE	9.54	Fitzgeral…
Manihot esculenta	ROOT	8.33	Fitzgeral…
Magnoliopsida	FRUIT	8.14	Fitzgeral…
Insecta	WHOLE	5.92	Fitzgeral…
Apoidea	WHOLE	5.13	Grzimek,B…
Aves	WHOLE	5.11	Grzimek,B…
Mammalia	WHOLE	4.47	Grzimek,B…
Ipomoea batatas	ROOT	4.17	Fitzgeral…
Apoidea	LARVAE	4.17	Fitzgeral…
Vertebrata	WHOLE	4.17	Fitzgeral…
Animalia	LARVAE	2.98	Fitzgeral…
Termitidae	EGGS	2.38	Fitzgeral…
Insecta	LARVAE	2.08	Nowak,R. …
Oligochaeta	WHOLE	1.92	Fitzgeral…
Cocos nucifera	LEAF	1.74	Nowak,R. …
Arecaceae	LEAF	1.67	Grzimek,B…
Oniscidea	WHOLE	1.19	Fitzgeral…
Gallus	WHOLE	1.04	Nowak,R. …
Cocos nucifera	SEED	0.76	Smith,A. …
Rodentia	WHOLE	0.69	Nowak,R. …
Cocos	LEAF	0.38	Smith,A. …
Magnoliopsida	NECTAR/JUICE	0.35	Nowak,R. …
Apoidea	EXUDATES	0.19	Smith,A. …

^a. Standardized values. See details for the calculations in Lintulaakso, K., Tatti, N. and Žliobaitė, I., 2022. Quantifying mammalian diets. Mammalian Biology, pp.1-15. [DOI]

Table 2. List of Standard Traits for Helarctos malayanus.
Trait	Selected Value	All Value(s)	Reference(s)
Dietary category: Eisenberg 1981	Frugivore-omnivore	Frugivore-Herbivore,Frugivore-omnivore	John L. G…
Dietary category: Miljutin 2009	Frugivore	Frugivore	John L. G…
Locomotion: Eisenberg 1981	Scansonal	Scansonal	John L. G…
Temporal habits	nocturnal	nocturnal	John L. G…

Table 3. List of Standard Measurements for Helarctos malayanus.
Measurement	Location	Unit	n Total	n Unknown	n Female	n Male	Min	Mean	Max	SD	Reference(s)
Body Mass (BM)	nan	g	-	-	-	-	-	57,076	-	-	Juhn, M.,…
Body Mass (BM)	nan	g	2	2	-	-	-	80,350	-	-	Engelman,…
Body Mass (BM)	nan	g	-	-	-	-	25,000	-	66,000	-	Freudenth…
Head-Body Length (HBL)	nan	mm	-	-	-	-	-	1,200	-	-	Engelman,…
Occipital condyle width (OCW)	nan	mm	2	2	-	-	-	51.85	-	-	Engelman,…

Table 4. List of other Measurements ^b. for Helarctos malayanus.
Measurement	Location	Unit	n Total	n Unknown	n Female	n Male	Min	Mean	Max	SD	Reference(s)
brainmass	nan	g	-	-	-	-	-	386	-	-	Engelman,…
skull_length	nan	mm	-	-	-	-	-	248	-	-	Engelman,…
femur circumference	nan	mm	-	-	-	-	-	65	-	-	Nelson, A…
humerus circumference	nan	mm	-	-	-	-	-	71	-	-	Nelson, A…
relative blade length (RBL)	nan	unitless	-	-	-	-	-	1	-	-	Juhn, M.S…

^b. other Measurements. Source Measurements which are not listed on or mapped with the Standard Traits..

Table 5. List of Data Sources for Helarctos malayanus.
Taxon	Reference	Match	Status
Helarctos malayanus	Gittleman,J. L., 1985, Carnivore body size: ecological and taxonomic correlates, Oecologia	1: Exact Taxon match	Verified - Accepted
Helarctos malayanus	Fitzgerald,C. S., 2002, Helarctos malayanus, Mammalian Species	1: Exact Taxon match	Verified - Accepted
Ursus malayanus	Nowak,R. M., 1991, Walker's mammals of the world,	50: Synonym	Verified - Accepted
Helarctos malayanus	Grzimek,B., 2003, Grzimek's Animal Life Encyclopedia: Mammals I-V,	1: Exact Taxon match	Verified - Accepted
Helarctos malayanus	Smith,A. T., 2010, A guide to the mammals of China,	1: Exact Taxon match	Verified - Accepted
Ursus malayanus	Nowak,R. M., 1991, Walker's mammals of the world,	50: Synonym	Verified - Accepted
Helarctos malayanus	Engelman, R.K., 2022. Occipital condyle width (OCW) is a highly accurate predictor of body mass in therian mammals. BMC biology, 20(1), p.37.	1: Exact Taxon match	Verified - Accepted
Helarctos malayanus	Nelson, A., Engelman, R.K. and Croft, D.A., 2023. How to weigh a fossil mammal? South American notoungulates as a case study for estimating body mass in extinct clades. Journal of Mammalian Evolution, 30(3), pp.773-809.	1: Exact Taxon match	Verified - Accepted
Helarctos malayanus	Freudenthal, M. and Martín-Suárez, E., 2013. Estimating body mass of fossil rodents. Scripta Geologica, 145, pp.1-513.	1: Exact Taxon match	Verified - Accepted
Ursus malayanus	Juhn, M.S., Balisi, M.A., Doughty, E.M., Friscia, A.R., Howenstine, A.O., Jacquemetton, C., Marcot, J., Nugen, S. and Van Valkenburgh, B., 2024. Cenozoic climate change and the evolution of North American mammalian predator ecomorphology. Paleobiology, pp.1-10.	50: Synonym	Verified - Accepted

Citation for this page: [The MammalBase community 2024. / CC BY 4.0. http://doi.org/10.5281/zenodo.7462864 Helarctos malayanus , Accessed 22nd November 2024 at https://mammalbase.net/me/14639]