MammalBase - Accepted taxon: Hyaena brunnea

Accepted taxon: Hyaena brunnea
Taxon:	Hyaena brunnea
Rank:	Species
Higher Classification:	CARNIVORA \| FELIFORMIA \| Hyaenidae \| Hyaena
Reference:	Wilson, Don E., and DeeAnn M. Reeder, eds. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), JHU Press, 2005.

Fig.1 - Ternary plot of the proximate analysis for the Hyaena brunnea diet.
CF (Crude Fibre) 4, ASH (Ash contents) 8, NFE (Nitrogen-Free-Extract) 17, CP (Crude Protein) 51, EE (Ether Extract) 20. See details for the calculations in Lintulaakso, K., Tatti, N. and Žliobaitė, I., 2022. Quantifying mammalian diets. Mammalian Biology, pp.1-15. [DOI]

Table 1. List of Diet Items for Hyaena brunnea.
TSN Complete name	Part	Ratio ^a.	Reference(s)
Mammalia	CARRION	16.44	Nowak,R. …
Otariidae	WHOLE	16.44	Grzimek,B…
Insecta	WHOLE	9.50	Damuth, J…
Vertebrata	CARRION	8.47	Damuth, J…
Magnoliopsida	FRUIT	7.63	Damuth, J…
Pedetes capensis	WHOLE	5.48	Grzimek,B…
Vertebrata	EGGS	4.73	Damuth, J…
Antidorcas marsupialis	WHOLE	4.38	Grzimek,B…
Rodentia	WHOLE	4.11	Nowak,R. …
Magnoliopsida	SHOOT	3.76	Damuth, J…
Otocyon megalotis	WHOLE	3.29	Grzimek,B…
Osteichthyes	WHOLE	2.37	Damuth, J…
Struthio camelus	EGGS	2.19	Grzimek,B…
Otididae	WHOLE	2.19	Grzimek,B…
Aves	WHOLE	2.09	Damuth, J…
Brachyura	CARRION	1.76	Nowak,R. …
Reptilia	WHOLE	1.49	Damuth, J…
Animalia	WHOLE	1.10	Grzimek,B…
Vertebrata	WHOLE	1.10	Grzimek,B…
Bovidae	WHOLE	0.60	Damuth, J…
Phocidae	WHOLE	0.59	Nowak,R. …
Panthera leo	WHOLE	0.30	Damuth, J…

^a. Standardized values. See details for the calculations in Lintulaakso, K., Tatti, N. and Žliobaitė, I., 2022. Quantifying mammalian diets. Mammalian Biology, pp.1-15. [DOI]

Table 2. List of Standard Traits for Hyaena brunnea.
Trait	Selected Value	All Value(s)	Reference(s)
Temporal habits	nocturnal	nocturnal	– John L. Gittleman, 1985, 'Carnivore body size: Ecological and taxonomic correlates', Oecologia, vol. 67, no. 4, pp. 540-554
Locomotion: Eisenberg 1981	Terrestrial	Terrestrial	– John L. Gittleman, 1985, 'Carnivore body size: Ecological and taxonomic correlates', Oecologia, vol. 67, no. 4, pp. 540-554
Dietary category: Miljutin 2009	Animalivore	Animalivore, Frugivore	– John L. Gittleman, 1985, 'Carnivore body size: Ecological and taxonomic correlates', Oecologia, vol. 67, no. 4, pp. 540-554, Juhn, M., Balisi, M., Doughty, E., Friscia, A., Howenstine, A., Jacquemetton, C., Marcot, J., Nugen, S., Van Valkenburgh, B. 2024. Cenozoic climate change and the evolution of North American mammalian predator ecomorphology. Paleobiology. None(None), pp.1-10. Available at: 10.1017/pab.2024.27.
Dietary category: Eisenberg 1981	Carnivore	Carnivore	– Juhn, M., Balisi, M., Doughty, E., Friscia, A., Howenstine, A., Jacquemetton, C., Marcot, J., Nugen, S., Van Valkenburgh, B. 2024. Cenozoic climate change and the evolution of North American mammalian predator ecomorphology. Paleobiology. None(None), pp.1-10. Available at: 10.1017/pab.2024.27.

Table 3. List of Standard Measurements for Hyaena brunnea.
Measurement	Location	Unit	n Total	n Unknown	n Female	n Male	Min	Mean	Max	SD	Reference(s)
Body Mass (BM)	nan	g	-	-	-	-	-	42,978	-	-	Juhn, M.,…
Body Mass (BM)	nan	g	8	-	-	8	-	47,100	-	-	Nelson, A…
Body Mass (BM)	nan	g	4	-	4	-	-	42,000	-	-	Nelson, A…
Body Mass (BM)	nan	g	-	-	-	-	26,900	-	73,000	-	Freudenth…
Head-Body Length (HBL)	nan	mm	8	-	-	8	-	1,230	-	-	Nelson, A…
Head-Body Length (HBL)	nan	mm	4	-	4	-	-	1,170	-	-	Nelson, A…

Table 4. List of other Measurements ^b. for Hyaena brunnea.
Measurement	Location	Unit	n Total	n Unknown	n Female	n Male	Min	Mean	Max	SD	Reference(s)
femur circumference	nan	mm	-	-	-	-	-	53	-	-	Nelson, A…
femur diameter	nan	mm	-	-	-	-	-	17	-	-	Nelson, A…
humerus circumference	nan	mm	-	-	-	-	-	58	-	-	Nelson, A…
humerus diameter	nan	mm	-	-	-	-	-	16	-	-	Nelson, A…
relative blade length (RBL)	nan	unitless	-	-	-	-	-	1	-	-	Juhn, M.S…

^b. other Measurements. Source Measurements which are not listed on or mapped with the Standard Traits..

Table 5. List of Data Sources for Hyaena brunnea.
Taxon	Reference	Match	Status
Hyaena brunnea	Gittleman,J. L., 1985, Carnivore body size: ecological and taxonomic correlates, Oecologia	1: Exact Taxon match	Verified - Accepted
Parahyaena brunnea	Damuth, J. 2010. ABSOLUT 6 a mammalian database compiled by the national center for ecological analysis and synthesis (NCEAS ) workshop on mammalian communities.	50: Synonym	Verified - Accepted
Hyaena brunnea	Nowak,R. M., 1991, Walker's mammals of the world,	1: Exact Taxon match	Verified - Accepted
Hyaena brunnea	Grzimek,B., 2003, Grzimek's Animal Life Encyclopedia: Mammals I-V,	1: Exact Taxon match	Verified - Accepted
Hyaena brunnea	Nowak,R. M., 1991, Walker's mammals of the world,	1: Exact Taxon match	Verified - Accepted
Parahyaena brunnea	Law, C.J., Blackwell, E.A., Curtis, A.A., Dickinson, E., Hartstone‐Rose, A. and Santana, S.E., 2022. Decoupled evolution of the cranium and mandible in carnivoran mammals. Evolution.	50: Synonym	Verified - Accepted
Hyaena brunnea	DeMers, A.C., 2023. Functional Significance of the Mandible, Tooth Roots, and Tooth Crowns, and their Implications for Fossil Dietary Inference (Doctoral dissertation, The Ohio State University).	1: Exact Taxon match	Verified - Accepted
Hyaena brunnea	Nelson, A., Engelman, R.K. and Croft, D.A., 2023. How to weigh a fossil mammal? South American notoungulates as a case study for estimating body mass in extinct clades. Journal of Mammalian Evolution, 30(3), pp.773-809.	1: Exact Taxon match	Verified - Accepted
Hyaena brunnea	Freudenthal, M. and Martín-Suárez, E., 2013. Estimating body mass of fossil rodents. Scripta Geologica, 145, pp.1-513.	1: Exact Taxon match	Verified - Accepted
Parahyaena brunnea	Juhn, M.S., Balisi, M.A., Doughty, E.M., Friscia, A.R., Howenstine, A.O., Jacquemetton, C., Marcot, J., Nugen, S. and Van Valkenburgh, B., 2024. Cenozoic climate change and the evolution of North American mammalian predator ecomorphology. Paleobiology, pp.1-10.	50: Synonym	Verified - Accepted
Parahyaena brunnea	Rowan, J., Beaudrot, L., Franklin, J., Reed, K.E., Smail, I.E., Zamora, A. and Kamilar, J.M., 2020. Geographically divergent evolutionary and ecological legacies shape mammal biodiversity in the global tropics and subtropics. Proceedings of the National Academy of Sciences, 117(3), pp.1559-1565.	50: Synonym	Verified - Accepted

Citation for this page: [The MammalBase community 2026. / CC BY 4.0. http://doi.org/10.5281/zenodo.7462864 Hyaena brunnea , Accessed 3rd May 2026 at https://mammalbase.net/me/14402]